[guided]The inverse branch is defined only on $\Lambda_2\times V$, so the solution must be defined on that same parameter neighbourhood. Define $u: \Lambda_2 \times V \to I$ by
\begin{align*}
u(\lambda,x)=Z(\lambda,s_\lambda(x),\theta_\lambda(x)).
\end{align*}
For each $\lambda\in\Lambda_2$, define $u_\lambda:V\to I$ by $u_\lambda(x)=u(\lambda,x)$. The map $Z$ is $C^k$ from the ODE smooth-dependence step, and the inverse branch $S(\lambda,x)=(s_\lambda(x),\theta_\lambda(x))$ is $C^k$ from the parameter-dependent inverse theorem. Therefore their composition is $C^k$, so $(\lambda,x)\mapsto u_\lambda(x)$ is $C^k$ on $\Lambda_2\times V$.
We now check the initial condition. Shrink $\Theta_2$ further, if necessary, so that $Y(\Theta_2)\subset V$. For $\theta\in\Theta_2$, the characteristic initial condition gives $X(\lambda,0,\theta)=x_\lambda(0,\theta)=Y(\theta)$. Since $S(\lambda,\cdot)$ is the chosen inverse branch of $X_\lambda$ on $V$, applying this inverse to $Y(\theta)$ gives $s_\lambda(Y(\theta))=0$ and $\theta_\lambda(Y(\theta))=\theta$. Hence $u_\lambda(Y(\theta))=Z(\lambda,0,\theta)=z_\lambda(0,\theta)=h(\lambda,\theta)$.
Finally we verify that $u_\lambda$ satisfies the first-order PDE. The defining identity is
\begin{align*}
u_\lambda(X(\lambda,s,\theta))=Z(\lambda,s,\theta),
\end{align*}
because $S$ is the inverse branch of $X_\lambda$ and $u_\lambda$ was defined by composing $Z$ with that inverse branch. Since $k\geq 1$, all functions in this identity are differentiable. Differentiating with respect to the characteristic time $s$ gives
\begin{align*}
\nabla u_\lambda(X(\lambda,s,\theta))\cdot \partial_s X(\lambda,s,\theta)=\partial_s Z(\lambda,s,\theta).
\end{align*}
The characteristic system identifies the two $s$-derivatives:
\begin{align*}
\partial_s X(\lambda,s,\theta)=a(X(\lambda,s,\theta),Z(\lambda,s,\theta))
\end{align*}
and
\begin{align*}
\partial_s Z(\lambda,s,\theta)=b(X(\lambda,s,\theta),Z(\lambda,s,\theta)).
\end{align*}
For an arbitrary $x\in V$, put $(s,\theta)=(s_\lambda(x),\theta_\lambda(x))$. The inverse identity gives $X(\lambda,s_\lambda(x),\theta_\lambda(x))=x$, and the definition of $u_\lambda$ gives $Z(\lambda,s_\lambda(x),\theta_\lambda(x))=u_\lambda(x)$. Substituting these two identities into the differentiated characteristic identity yields
\begin{align*}
a(x,u_\lambda(x))\cdot \nabla u_\lambda(x)=b(x,u_\lambda(x)).
\end{align*}
Thus $u_\lambda$ is a classical solution, it has the prescribed initial data, and the solution family depends $C^k$ on $(\lambda,x)$.[/guided]